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Anomalipes zhaoi
Topic Started: Jun 12 2018, 04:21 PM (80 Views)
Taipan
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Anomalipes zhaoi

Temporal range: Late Cretaceous, Campanian

Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Order: Saurischia
Suborder: Theropoda
Family: †Caenagnathidae
Genus: †Anomalipes Yu et al., 2018
Type Species:Anomalipes zhaoi Yu et al., 2018

Anomalipes zhaoi ("meaning Zhao Xijin's unusual foot") is an extinct caenagnathid dinosaur discovered in the Cenomanian-aged Wangshi Group in China. It is the only species in the genus Anomalipes.

Discovery
The remains of Anomalipes were discovered within a Shantungosaurus bonebed at the Kugou locality. The holotype (ZCDM V0020, housed at Zhucheng Dinosaur Museum, Zhucheng, Shandong) is an incomplete left hind-limb, including a partial left thigh, shin and shank, a complete metatarsal III and two toe bones.

Etymology
Anomalipes is derived from the Latin anomalus (peculiar, abnormal, not easily classified) and pes (foot), referring to the unusual shape of its foot. The species epithet, zhaoi, was named in honour of Zhao Xijin.

Posted Image
Figure 5 Simplified oviraptorosaurian phylogenetic tree, showing size ranges for basal oviraptorosaurs, the Caenagnathidae, and the Oviraptoridae. Grey boxes represent body mass ranges for three oviraptorosaurian groups: basal oviraptorosaurs, oviraptorids, and caenagnathids. See the electronic supplementary material for estimated body masses of various oviraptorosaurian species.
Abbreviations: Caud: Caudipteryx zoui; Avim: Avimimus portentosus; Conc: Conchoraptor gracilis; Micr: Microvenator celer; Wula: Wulatelong gobiensis; Citi: Citipati osmolskae; Nank: Nankangia jiangxiensis; Anom: Anomalipes zhaoi; Neme: Nemegtia barsboldi; Anzu: Anzu wyliei; Giga: Gigantoraptor erlianensis.


Holotype:
ZCDM V0020 (Zhucheng Dinosaur Museum, Zhucheng, Shandong, China), an incomplete left hindlimb, including the left femur missing the distal end, the left tibia missing the proximal end, the left fibula missing the distal and proximal ends, a complete metatarsal III and two pedal phalanges. Although these bones are disarticulated, they are inferred to be derived from a single theropod individual given that 1) they were preserved in a small area of less than 0.3 square metres within a Shantungosaurus bonebed; and 2) no other theropod skeletal elements are preserved nearby.

Locality and horizon:
Kugou, Zhucheng City, Shandong Province, China. Upper Cretaceous Wangshi Group.

Diagnosis:
A new caenagnathid with the following unique combination of features: femoral head anteroposteriorly narrow and somewhat deflected posteriorly; accessory trochanter low; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior flange near proximal end, medial hemicondyle much narrower than lateral hemicondyle, and longitudinal groove on distal articular surface; and pedal phalanx II-3 with lateral collateral groove deeper and more dorsally located than medial groove.




Journal Reference:
Yilun Yu, Kebai Wang, Shuqing Chen, Corwin Sullivan, Shuo Wang, Peiye Wang and Xing Xu. 2018. A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Comments on Size Variation Among Oviraptorosaurs. Scientific Reports. volume 8, Article number: 5030. DOI: 10.1038/s41598-018-23252-2

Abstract
The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen’s systematic position was assessed by three numerical cladistic analyses based on recently published theropod phylogenetic datasets, with the inclusion of several new characters. Anomalipes zhaoi differs from other known caenagnathids in having a unique combination of features: femoral head anteroposteriorly narrow and with significant posterior orientation; accessory trochanter low and confluent with lesser trochanter; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior flange near proximal end, highly asymmetrical hemicondyles, and longitudinal groove on distal articular surface; and ungual of pedal digit II with lateral collateral groove deeper and more dorsally located than medial groove. The holotype of Anomalipes zhaoi is smaller than is typical for Caenagnathidae but larger than is typical for the other major oviraptorosaurian subclade, Oviraptoridae. Size comparisons among oviraptorisaurians show that the Caenagnathidae vary much more widely in size than the Oviraptoridae.

https://www.nature.com/articles/s41598-018-23252-2
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